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HNA Summary Report
The Importance of Scale in Large River Ecosystems

Depending on their mobility and life requirements, the scale or geographic extent of habitats is important to river organisms. Aquatic and floodplain species in the UMRS have adapted to the size of river habitats and the dynamic set of river habitat conditions for millennia. The major landforms of the present UMRS developed over 11,000 years ago during the retreat of glaciers. The north-south orientation of the Mississippi River provided refuge for species during glacial times and continues to provide a corridor for migration and dispersal of many life forms.

Basin and Continental Scales
The basin and larger scales are appropriate when considering the habitat needs of animals that migrate over long distances. Among fish, paddlefish, sturgeon, skipjack herring, and the American eel are notable long distance migrants. Many bird species migrate between North, Central, and South America. Although many species migrate beyond the UMRS, they all require specific habitat resources when they use areas along the rivers.

1.Place the cursor over the image at lower left for a preview of the individual reaches.
2.Click within the image at lower left for a detailed view of the land cover/use for each reach.
UMRS basin
UMRS reach

System Scale
The UMRS, as defined by EMP authorizing legislation, includes the Upper Mississippi River from Minneapolis, Minnesota to Cairo, Illinois, the entire Illinois River, and navigable portions of the Minnesota, St. Croix, Black, and Kaskaskia Rivers. This HNA covers the aquatic and floodplain areas of the UMRS.

River Reach Scale
Twelve river reaches (see image above) have been defined according to the dominant geomorphic features of the Mississippi and Illinois Rivers. These geomorphic features provide the template upon which plant communities and animal habitats developed. The geomorphic features (see sidebar) of a river reach also influence the river’s response to impoundment for navigation. The shallow reservoirs, or navigation pools, in some river reaches developed broad, open-water impounded areas, whereas others show little apparent plan form (see sidebar below) change due to impoundment. Habitats and the ecological communities they support differ among river reaches, thus resource opportunities, problems, and management differ among the river reaches.

Geomorphology is the geological study of the configuration and evolution of land forms. Fluvial geomorphology is the study of the development of land forms, streams, and rivers under processes associated with running water. Plan form is the shape of a landscape as seen from above, or in map view. The GIS maps used throughout this report are plan form views of habitat. Plan form images from different time periods are used to measure change in the river system.

Lake sturgeon
Lake sturgeon and other fish species may migrate hundreds of miles among river reaches.

Some species migrate over a regional scale to complete their life cycle. Regional migratory fish include species such as walleye, smallmouth bass, white bass, and some sucker species that move upstream to spawn, often in tributaries to the mainstem rivers. Dams and tributary habitat degradation have reduced access to habitat for these fishes. Many bird species migrate along the Mississippi River to find warmer winter temperatures in southern states.

Navigation Pool Scale
Bellevue Lock and DamThe mainstem dams of the UMRS navigation project formed a series of shallow reservoirs called navigation pools. The pool scale is important in assessing the physical environment that defines habitat for species that display seasonal movements of about 20 miles or less. UMRS navigation pools differ in their mix of habitats among river reaches. Wide-ranging residents within navigation pools include a variety of fishes such as largemouth bass, northern pike, catfish, and buffalo which make seasonal movements to find appropriate habitats to spawn, feed, or over-winter. Some species require flooded vegetation to spawn, others need structure and undercut banks, and some require firm bottom substrate. Most fish species require winter habitat with low current, adequate levels of dissolved oxygen, and water temperatures higher than freezing. All seasonal habitat needs for resident species cardinalmust be met within a navigation pool because dams restrict access to other navigation pools. Resident birds maintain home ranges over many square miles. Cardinals, blue jays, woodpeckers, crows, and many others may use both floodplain and upland habitats. Some bird species may nest in one floodplain habitat and feed in another which requires that important habitats are available within their home range.

Habitat Scale
The habitat scale is the level that is actually occupied by organisms. UMRS habitats must provide suitable resources to meet the needs of a variety of riverine organisms. Habitats for long-distance migrants and wide-ranging species are large, while relatively immobile organisms such as freshwater mussels have small habitat areas. Most riverine organisms have habitat needs that can be measured in square yards to tens of acres. Many river organisms require diverse habitat conditions, with multiple habitat types in close proximity. Most river processes act at the habitat scale and protection and restoration is generally focused at this scale.

crappie
Crappie, bluegills, and many minnows may live their entire life in one backwater lake.

Many animal species have small home ranges that meet all their life history needs. Even migratory species will use small home ranges within their seasonal habitats. For species with system-wide distribution it is important that criticalmuskrat habitats are available and of suitable quality to support local populations. Aquatic invertebrates are generally restricted to small areas, but may drift in currents or migrate during adult aerial stages. Freshwater mussels are a particularly threatened group of animals that have suffered greatly through harvest or pollution due to their lack of mobility. Many panfish and minnows and most small mammals have limited ranges.

 
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