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We defined landscape composition variables as those variables calculated solely from the proportion or amount of different land use/land cover classes, without any information regarding the spatial arrangement of land cover patches.
We defined landscape configuration variables as those variables calculated from information regarding the spatial arrangement of patches, where a patch is defined as a homogenous area of a single land use/land cover class.

References
Fauth, P. T., E. J. Gustafson, and K. N. Rabenold. 2000. Using landscape
metrics to model source habitat for Neotropical migrants in the
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Gustafson, E. J. 1998. Quantifying landscape spatial pattern: what is the state of the art? Ecosystems1:143-156.
Gustafson, E. J., and G. R. Parker. 1992. Relationships between landcover proportion and indices of landscape spatial pattern. Landscape Ecology 7:101-110.
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Topography, the physical and natural features of a landscape, is
known to affect distribution of plants and thus the birds that use
them for cover, food, nesting, and other life history requirements.
However, rarely are topographic considerations included in habitat
modeling. When topography is considered, usually this consideration
is limited to elevation, slope, and aspect. Ruggedness of terrain
and slope position, two other areas of topography, are important
because they influence microclimate, cover from predation, and susceptibility
to disturbance by humans. These additional areas we consider in
our current modeling exercise.
References
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Gustafson, E. J., N. L. Murphy, and T. R. Crow. 2001. Using a GIS model to assess terrestrial salamander response to alternative forest management plans. Journal of Environmental Management 63:281-292.
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Renfrew, R. B., and C. A. Ribic. 2002. Influence of topography on density of grassland passerines in pastures. American Midland Naturalist 147:315-325.
Riley, S. J., S. D. DeGloria, and R. Elliot. 1999. A terrain ruggedness index that quantifies topographic heterogeneity. Intermountain Journal of Sciences 5:23-27.
Samways, M. J. 1990. Land forms and winter habitat refugia in the conservation of montane grasshoppers in southern Africa. Conservation Biology 4:375-382.
Swanson, F. J., T. K. Kratz, N. Craine, and R. G. Woodmansee. 1988. Landform effects on ecosystem patterns and processes. BioScience 38:92-98.
Climate is one of the fundamental determinants of a species range. For many species, species habitat exists beyond the usual distribution This habitat would be suitable for occupancy but for the deleterious effects of climate.
Climate
data was supplied by the Canadian Forest Service. They derived
spatially-explicit climate data layers for North America from thin
plate spline smoothing algorithms applied to weather station
data. Their approach accommodated local spatial autocorrelation
and topographic effects.
References
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and C. J. Ralph (eds). Wildlife 2000: Modeling habitat relationships
of terrestrial vertebrates.
Parmesan, C., T. L. Root, and M. R. Willig. 2000. Impacts of extreme weather and climate on terrestrial biota. Bulletin of the American Meteorological Society 81:443-450.
Porter, W. P., S. Budaraju, W. E. Stewart, N. Ramankutty. 2000. Calculating climate effects on birds and mammals: Impacts on biodiversity, conservation, population parameters, and global community structure. American Zoologist 40: 597-630.
Root, T. L. 1988. Environmental factors associated with avian distributional limits. Journal of Biogeography 15:489-505.
Thomas, C. D., and J. J. Lennon. 1999. Birds extend their ranges northward. Nature 399:213.
Venier, L. A., D. W. McKenney, Y. Wang, J. McKee. 1999. Models of large-scale breeding-bird distribution as a function of macro-climate in Ontario, Canada. Journal of Biogeography 26:315-328.
Watson, A., R. Moss, and P. Rothery. 2000. Weather
and synchrony in 10-year population cycles of Rock Ptarmigan and
Red Grouse in Scotland. Ecology 81:2126-2136.